This species is distinguished by the following characters: monospondylous vertebrae 46-52; precaudal total vertebrae 124-131; posterior nasal flap developed; first dorsal fin origin anterior to free rear tips of pectoral fin; snout length short with snout tip to oral length < 1.8 times internarial width; embryos and juveniles with dark complicated lines and rings; no accessory cartilage at symphysial part of upper jaw (Ref. 55581).